After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise ...

The San Diego Zoo announced Gramma, the Galápagos tortoise, died on Nov. 20. She was about 141 years old based on estimates ...

Gramma arrived at the San Diego Zoo sometime between 1928 and 1931 and was supposedly several decades old at the time.

Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died at an estimated 141 years of age.

Gramma the tortoise, believed to be 141 years old and the oldest resident of the San Diego Zoo, was euthanized after ...